Nightmare of Nunnally starts with Lelouch walking in the Shinjuku ghetto, after having defeated a noble in chess and leaving behind Rivals, he encountered a runaway truck being crashed and proceeds in helping the driver of the truck. He asks the injured driver if he's alright and replied about the cargo, were a capsule has fallen out and opens, revealing C.C. and a mysterious doll, Nemo, but later he encountered a group of Sutherlands preparing to attack but retreats in the last minute. When Lelouch questions why, the driver revealed that the girl is not an ordinary human and was later shocked when the driver revealed that the Purity League plans to destroy the ghetto. Lelouch run away with C.C. after the driver apologizes for getting him involved, but was caught by a falling debris, which forced both C.C. and Nemo to activate their powers before the area they are in was destroyed.
After the Purity League and Clovis's forces were defeated, Lelouch reappeared in the Governor General's office as Zero (in a suit of armour) and was unharmed as Clovis fired his gun at him, but Lelouch proceeds in killing Clovis and written with his blood the Kanji words "Tenchu" on the wall. He later made contact with Ohgi's resistance group and propose an allegiance. After the Lake Kawaguchi incident, Lelouch saved most of the hostages and Kallen from a rogue group of the Japanese Liberation Front and killed the groups leader, Kusakabe, while a Knightmare battle involving the Mark Nemo defeating both the Brittanian and Japanese forces and battling one of the Irregulars Alpha units was in progress. He later revealed to the media his own version of the Black Knights and declares war on Britannia.
In the battle against Brittanian's forces in the Saitama ghetto, Lelouch confronts Cornelia while easily shutting down both Darlton and Guilford's Knightmare's power supply and used his ability to force her out from her Knightmare, completely injuring her. He demanded information from her of the Emperor and his knights, which Cornelia finds amusing for him to oppose the Emperor himself but refused by trying to kill herself which Lelouch was about to prevent her. However he was confronted by Suzaku in his Lancelot Knightmare and kicked him on to a truck but survived the attack, much to the shock from Cornelia, and the two battled each other. Lelouch fought against Suzaku, with the same fighting style as his, but was surprised that his abilities was negated and wonders if he's a Geass user and prepares to release his full power, however he retreated when he sensed Nunnally in trouble. He manages to arrived on time to prevent the Irregulars' attack and teleported with her in the Mark Nemo. He later revealed himself to Nunnally and told her not to abuse the power, but also revealed that he is Zero and demands that Nunnally that she no longer fight with Mark Nemo before giving her a paper crane and leaves.
Later at the Battle of Narita, Lelouch begins his operation against Cornelia's forces, who had already begun its assault against the JLF base, leaving her exposed for the Black Knights and wiped out half of her forces using the Avalanche strategy. Lelouch late took down Guilford and confronted Cornelia. Lelouch begun his assault using his D.O.M. Hostia ability, but Cornelia manage to avoided and fired missiles at him, but has completely underestimated him and Lelouch has stopped them instantly and quickly defeated her from behind. As he prepares to finish her off, Lelouch was once again interrupted by Suzaku in his Lancelot. The two both battled and Lelouch questioned Suzaku about his power as a Wired and was not satisfied, which Suzaku did not understand, leaving Lelouch to use his ability and reveal to him the history of his power involving Jeanne D'Arc. He also revealed to Suzaku that he has made a contract with C.C. and became Zero, which has shocked Suzaku, and offers him to be his ally so they can build a gentler world for Nunnally. However Suzaku refused, believing him to be the witch C.C. in the form of Lelouch and attacks him in rage. Lelouch was surprised that Suzaku's power had increased since their last battle and had managed to defeat him, but was interrupted by the arrival of Mark Nemo, who he warns her to get away. Later an unknown unit fired a Hadron Cannon on the battle field, in the middle of a battle between Nunnally's Mark Nemo and Alice's Alpha, and Lelouch suspect that it is his younger brother, Rolo Vi Britannia.
After the battle of Narita, Lelouch went to the Governor General building and appeared in his normal form to meet with Suzaku, who brought out his gun and sword at him, and revealed to him that the pilot of Mark Nemo who is in custody of "The Eden Vital Religious Order" is Nunnally, which has completely shocked him in disbelief, and request to him that they both must rescue her from execution, which he now agrees and pass the information to Euphemia. Both Lelouch and Suzaku arrived in time to prevent Nunnally's execution and manage to save her, but are confronted by both Rolo and Anya in their Knightmare Frames. Lelouch fought against Rolo, but was surprised that he used Quantum Shift to summon his Knightmare Frame, Vincent, and revealed himself as the Demon King before entering in his Knightmare. This forced Lelouch to use his Quantum Shift to summon his Knightmare, Gawain, and fought. Lelouch fired his Hadron Cannon at him, but Rolo used his Geass to destroy his Gawain and witness Suzaku being defeated by Anya. They later saw Alice, who Lelouch knows that she made a contract with Nemo and witness her summon her Knightmare, Code Geass, and defeated Anya in the process. He later escaped after Alice used her Knightmare's power to bring down the entire facility, knowing Nunnally being safe with her, and manage to save Suzaku.
After the Emperor, Charles zi Britannia, declares the Holy Nation of Eden Vital, Lelouch ordered the Black Knights to break free Suzaku and Euphemia, with Cornelia, Guilford, Lloyd, and Cecile, with Kaguya and the members of the Kyoto House to continue their negotiation. With the two sides coming to an agreement with the condition that Euphemia becomes the new ruler of Britannia, Lelouch orders the Black Knights to fight along side Britannia, against Charles and the Holy Nation of Eden Vital. When both forces of the Black Knights and Britannnia head towards Kamineji Island, Lelouch and Suzaku both fought against an army of zombified Knightmares and the undead Knight of the Rounds, until they all died from the Emperor's defeat.
After the Emperor's defeat and Zero's death was announced, Lelouch appeared in front of Suzaku and Euphemia to bid them farewell as it is his last day as Lelouch. He then meets with Nunnally, who is now a goodwill ambassador of Japan, and discuss about their Geass. Lelouch then discuss that from Nunally's Geass, "The Zero", he had been freed from C.C.'s duties by death, as her soul now returned to Eden Vital and has been made as her successor like many others before her and all the Geass that she bestowed upon have now disappeared. They also discuss about Nunnally's vision's of alternative realities, one reality of which is the event of the anime, and was furious that she did not accept their mother and father's plan, but acknowledges her dream for a peaceful world, to which he gave her a hug and bids farewell before saying that her brother will still love her. At the conclusion of the story, Lelouch inherits C.C.'s name and immortality, becoming C.C. The Demon King and goes forth to spread Geass and to promote conflict around the world.

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