In an alternative event of R2, Lelouch ordered the Black Knights to take one million Japanese and escapes to the Chinese Federation. However a group of Britannian soldiers are in pursuit, and plans a surprise attack on them. Later in an alternative event of episode 11, Lelouch as Zero was in a middle of a battle against the Chinese Federation for kidnapping Tianzi and won after exposing the eunuches plot and had them killed in their command ship. They are later confronted by the Britannian military forces led by Schneizel, along with the Knight of Rounds, Suzaku, Gino, Anya, and Luciano in their respected Knightmares. As the two sides prepare to fight each other an unknown black hole suddenly appeared and sucked all sides in, except for the Black Knight's flagship, Ikaruga, and a Britannian carriership with Schneizel inside, who both manage to escape.
Lelouch, along with Kallen and C.C., have arrived in Planet and suddenly appeared in a desert, were they witness a traces of a bonfire being spoted a ruined city. The group later entered a ruined dome, where they were approached by an unknown units, later revealing themselves to be Sousuke Sagara, Melissa Mao, and Kurz Weber from the Full Metal Panic! series in their respected Arm Slave. The two side begun attacking each other, believing each other to be the enemy, until a group of SPR Icon units appeared and starts attacking them, causing both sides to work together and successfully defeated them. They later meet a second group consists of mechs from both Super Dimension Century Orguss and Overman King Gainer, with Olson D. Verne in his Orguss II unit speaking on behalf to join forces, in exchange for information. Lelouch who is also speaking on behalf of his group accepted his offer.
As the group come to a truce and decide to work together, they later encountered Suzaku, in his Lancelot, along with the Avalon. Lelouch tried to convince him that they are not here to fight and asked to cooperate with them, but Suzaku refused and prepared to attack, but was interrupted by the arrival of a group of SPR forces and a defense arms. The girl in the defense arm introduces herself as Spring One and offers to guide them through this world because she feels sorry for them. However she is easily seen through by Zero, and he rejects it. In reply, Spring begun the attack on Zero's group, while the group was ordered by Zero to defend the Avalon. As a large number of SPR fleet arrived after Spring's escape, the SMS Macross Quarter from the Macross Frontier series arrived, along with the Ark Alpha, and wiped out all of the remaining fleet. Lelouch and the group were surprised by its transformation and was contacted by Jeffrey Wilder, Captain of the Macross Quarter, confirming Lelouch's suspicion as their allies and decided to join them with the group, along with Suzaku who also agreed.
Lelouch and his group had eventually side with Autumn Four and boarded on her ship, were he had met with Suzaku, along with Lloyd and Cecile, who had also side with Autumn, leading both to had a small quarrel but eventually put their differences aside. Later in the briefing room, after Autumn confessed to memory loss, he met with Autumn alone and revealed himself as Lelouch to her and used Geass on her to unlock her memories for answers, but was shocked when it did not have any effect on her. Autumn, aware of what he was doing, tells him that he and the others will be ready for their next mission and left. Lelouch has started to become fascinated with Autumn as his Geass did not work on her. Later in one of the missions, Lelouch and the others fought against Spring One in her Core unit, Sea Plant, which Autumn later delivered the final blow in her Core unit, Alpha Heart, causing Spring to escape. With many of the group starting to have doubts about Autumn, Lelouch question if she is really human, which she confirmed is true - that she was an android.
Later the Ark Alpha crew arrived at a Geo Plant facility station, and Lelouch was surprised by the appearance of Bonta-Kun, which took down two large Icon's easily. After passing through different levels, they encountered Cynthia Lane of Trans-Siberian Railway is blocked with Asuham Boone in front of them, from Overman King Gainer, and Zero notice Cynthia has lost the will to fight, leading Asuham Boone to attack and was defeated. This led Asuham to retreat, and the crew let Gainer convinced Cynthia to join them.
Crew of Ark Alpha succeed in the invasion to the centre of Geo Plant. However, Anti-U.N. forces and Summer One waited for them there. Summer One speaks the reason why there is no human in planet Area to them before the fight. Autumn believes that Ark Alpha is the hope of mankind, but Summer confirms that the humans have been destroyed at the Seasons hands, which has shocked the rest of the Ark-Alpha crew and begun attacking him in his Geo Breaker. As Summer One made his escape after his defeat and the collapse of the Geo Plant, Autumn went and secures data from Geo Breaker and request everyone to escape, believing that it would help them. Impressed with her determination, Lelouch assisted her and protected her from the falling debris with the others clear a path for her escape.
Near the end of the game, the rest of the group fought once more against the first three Seasons in their respective Plants. Winter then activated the ACE system, but was surprised that it would not work, and the group were later surprised as Winter was taken over an unknown entity, who revealed himself as Doctor Shiki, whom they then fought in Winter's Core unit, Four Seasons. As the group manage to defeat him, Doctor Shiki escape and both Autumn Four and Lelouch went after him. As Autumn was about to kill him, Doctor Shiki revealed that his death will destroy Eria, which Lelouch then confronts him, thinking his death would save Planet Area, and realises Doctor Shiki's mind is connected to Winter's artificial brain. Lelouch then used his Geass on Doctor Shiki to suffer an eternal nightmare, causing him to suffer an immediate breakdown and collapsed, and escape with Autumn Four. System ACE was later activated and Autumn Four suppressed the black hole being expanded and opens the gate, leading the group to return to their own world. As Lelouch and the others return to their world in the Chinese Federation, he request to Suzaku that the two would speak. The two had a brief conversation and went to there separate ways, with Lelouch knowing that they will be enemies again.
Lelouch is set to appear in the spin-off with his Shinkiro.
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